RT R T 1 16 q_Na_o c_Nae W_e q_Na_i c_Nai W_i q_K_o c_Ke W_e q_K_i c_Ki W_i q_Pi c_P_i W_i q_ADP c_MgADP W_i q_ATP c_MgATP W_i q_H 10 pH 1000 W_i z_2 1 z_1 t q_1 v_4 v_1 t q_2 v_1 v_2 t q_3 v_2 v_3 t q_4 v_3 v_4 u_e q_mem C_m Q_1 K_1 q_1 Q_2 K_2 q_2 Q_3 K_3 q_3 Q_4 K_4 q_4 Q_Na_o K_Nae q_Na_o Q_Na_i K_Nai q_Na_i Q_K_o K_Ke q_K_o Q_K_i K_Ki q_K_i Q_ATP K_ATP q_ATP Q_ADP K_ADP q_ADP Q_Pi K_Pi q_Pi Q_H K_H q_H v_1 kappa_1 Q_1 Q_K_o 2 Q_2 Q_Pi Q_H v_2 kappa_2 Q_2 Q_ATP Q_3 Q_K_i 2 v_3 kappa_3 Q_3 Q_Na_i 3 Q_4 Q_ADP z_1 F u_e RT v_4 kappa_4 Q_4 Q_1 Q_Na_o 3 z_2 F u_e RT q_tot q_1 q_2 q_3 q_4 Adenosine_tot q_ATP q_ADP q_3 Phosphate_tot 3 q_ATP 2 q_ADP q_Pi q_1 3 q_3 q_4 lambda_1_NKE kappa_2 kappa_4 lambda_2_NKE K_1 K_2 lambda_3_NKE K_1 K_3 lambda_4_NKE K_1 K_4 e_0 F u_e RT A_0_NKE q_tot kappa_2 K_1 A_1_NKE Q_Na_i Q_Na_o 3 Q_ADP Q_Pi Q_ATP Q_K_i Q_K_o 2 e_0 A_2_NKE Q_Pi Q_ATP Q_K_o 2 lambda_2_NKE Q_ATP A_3_NKE Q_ADP Q_Pi Q_ATP Q_K_i Q_K_o 2 lambda_2_NKE Q_ADP Q_K_i 2 Q_ATP lambda_3_NKE Q_ADP lambda_4_NKE Q_Na_i 3 z_1 F u_e RT A_4_NKE lambda_1_NKE z_1 F u_e RT Q_Na_o 3 Q_Pi Q_ATP Q_K_o 2 e_0 AA_NKE A_1_NKE A_2_NKE A_3_NKE A_4_NKE v_NKE A_0_NKE AA_NKE Q_Na_i Q_Na_o 3 Q_ADP Q_Pi Q_ATP Q_K_i Q_K_o 2 e_0 u_1 RT Q_1 u_2 RT Q_2 u_3 RT Q_3 u_4 RT Q_4 u_K_o RT Q_K_o u_K_i RT Q_K_i u_Na_o RT Q_Na_o u_Na_i RT Q_Na_i u_ATP RT Q_ATP u_ADP RT Q_ADP u_Pi RT Q_Pi u_f1 u_1 2 u_K_o u_r1 u_2 u_Pi u_f2 u_2 u_ATP u_r2 u_3 2 u_K_i u_f3 u_3 3 u_Na_i u_r3 u_4 u_ADP z_1 F u_e u_f4 u_4 u_r4 u_1 3 u_Na_o z_2 F u_e Power_external 2 u_K_o u_Pi v_1 u_ATP 2 u_K_i v_2 3 u_Na_i u_ADP z_1 F u_e v_3 3 u_Na_o z_2 F u_e v_4 Power_dissipation u_f1 u_r1 v_1 u_f2 u_r2 v_2 u_f3 u_r3 v_3 u_f4 u_r4 v_4